When the head rotates down, the eyes turn up.
If the VOR where perfect (and could rotate a full 90 deg.) the eye's would remain still in space and now be pointing up in the head.

Copywrite © 1996 Tutis Vilis and Douglas Tweed
Department of Physiology
University of Western Ontario
London Ontario Canada
Updated October 22, 1996

Comments welcome. Email to tvilis@physiology.uwo.ca